Fig. 12. All variable nucleotide sites found in ITS sequences among P. lactiflora, P. mairei. and P. obovata. Paeonia russi shows additivity at all the sites except site 8 that are variable between P. lactiflora and P. mairei. suggesting that P. russi is derived via hybridization between these two species. Paeonia cambessedesii has only partial additivity, and its origin is discussed in the text. The site numbers (1-12) corresponding to the sites numbered separately for ITS 1 and ITS 2 from 5' to 3': ITS 1: 49, 108, 131, 137, 138, 169, 226, 227; ITS2: 31, 38, 82, 179.